The locations of the eight forest and two oil palm plantation study areas in Sabah, Malaysian Borneo.

<p>Numbered polygons represent the different study areas, defined by the locations of the outermost camera stations: 1. Crocker Range Park; 2. Malua Forest Reserve; 3. Danum Palm Plantation; 4. Danum Valley Conservation Area (surveyed on two separate occasions); 5. Ulu Segama Forest Reserve; 6. Minat Teguh plantation; 7. Kabili-Sepilok Forest Reserve; 8. Lower Kinabatangan Wildlife Sanctuary; 9. Tabin Wildlife Reserve; 10. Tawau Hills Park (surveyed on two separate occasions). Inset shows the island of Borneo. Land use data from 2010 [<a href="http://www.plosone.org/article/info:doi/10.1371/journal.pone.0200828#pone.0200828.ref036" target="_blank">36</a>].