Sample collection sites in Sabah, Malaysian Borneo, Southeast Asia.

<p>(a) Map with Borneo, Southeast Asia, and the position of sampling sites framed. (b) Enlarged map with Sabah, Malaysian Borneo, and the position of the two sampling sites: the LKWS with Lots 1–10 (dark grey) along the Kinabatangan River (black line) with DGFC (white house) in Lot 6, and the LBPMS near the coastline (grey house). LKWS = Lower Kinabatangan Wildlife Sanctuary, LBPMS = Labuk Bay Proboscis Monkey Sanctuary, DGFC = Danau Girang Field Centre.