Study area.

<p>Map showing the location of Sabah (inset) on the island of Borneo in South East Asia, and the distribution of three forest reserves with different post-logging regeneration ages. East Sabah: Tabin Wildlife Reserve (TWR). Central Sabah: Malua Forest Reserve (MFR). South central Sabah: Maliau Basin Conservation Area Buffer Zone (MBCABZ) with a small portion of Gunung Rara Forest Reserve directly to the south of MBCABZ). Maps were generated using ArcGIS® software version 10.1 by ESRI, with data from Natural Earth and the Sabah Forestry Department.