Atherhagiox Grimaldi 2016, new genus

Atherhagiox, new genus DIAGNOSIS: Differs from living and extinct athericids by having base of fork of R 4+5 significantly proximal to level of apex of cell d (i.e., cell d is quite long). Further, veins M 1 and M 2 sinuous as in Galloatherix; like this genus and Sinocretomyia and Palaepangonius, R 2+3 also sinuous, but its apex not in contact with apex of R 1. Atherhagiox further differs from Sinocretomyia by the reniform (vs. ovoid) basal flagellomere. TYPE SPECIES: Atherhagiox ambiguum, new species. ETYMOLOGY: The generic name is a combination of the two names of type genera of Athericidae and Rhagionidae — Atherix and Rhagio —and refers to the hybrid set of characters of the new genus. The gender of the name is neuter. COMMENTS: Atherhagiox have similarities with certain Rhagionidae, and the boundaries between this family and Athericidae have historically been vague. Like the rhagionid genus Symphoromyia, the new genus has a reniform basal flagellomere with an aristalike terminal section of the flagellum and a sinuous R 2+3; both genera also have 0-2-1 tibial spurs. Other similarities are plesiomorphic: a vein R 2+3 that does not converge with or approximate the apex of R 1; suprametacoxal pit (found in all athericids except Dasyomma) is lacking; cerci 2-segmented, and female terminalia telescoping. However, like athericids and unlike rhagionids, Atherhagiox have a deeply excised, lappetlike basicosta and arched palps. Mandibles are present in the female (these are absent in most Rhagionidae with the exception of Symphoromyia).Published as part of Grimaldi, David A., 2016, Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) In Amber From The Cretaceous Of Myanmar: Brachycera In Cretaceous Amber, Part Vii David A. Grimaldi, pp. 1-132 in Bulletin of the American Museum of Natural History 2016 (408) on pages 35-36, DOI: 10.1206/0003-0090-408.1.1, http://zenodo.org/record/538094