Izleiina BLAGODEROV & GRIMALDI 2004, new genus

Izleiina, new genus DIAGNOSIS: Facets round. Lateral ocelli not touching eye margins. Clypeus setose. Sc meets C just beyond level of base of M3; Sc 2 absent. Length of R 1 2Χ that of r­m. Section M3, and fork of M 1 and M 2 –M 3+4 reduced; CuA weakened. M interrupted at base. 3+4 TYPE SPECIES: Izleiina mirifica, n.sp. ETYMOLOGY: Name is derived from the tribe Leiini. COMMENTS: We assume that in the new ge­ nus vein M 1 is entirely reduced. As a rule, in most leiine genera, vein M 1 reaches the wing margin at or even before the wing apex. They have retained a complete vein that is homologous to CuA based on the length and shape of the vein, and which is strongly curved back and ends at the level of RS1, as in most Leiini. The new genus differs from Neoclastobasis Ostroverchova by the lateral ocelli touching eye margins, and from Clastobasis Skuse and Neoclastobasis by C ending beyond the tip of R 5, and by the presence of the base of M 3+4. Having R 1 relatively long and Sc ending at C makes the genus similar to Rondaniella Johannsen and Indoleia Edwards, but differs from them by reduction of the veins in the medial sector. Such a reduction also occurs in other leiine genera, for example Novakia Strobl and Sigmoleia Tonnoir and Edwards, but they have Sc very short and free. Species of Cycloneura Marshall and Paracycloneura Tonnoir and Edwards also have M unbranched and M 3+4 free or reduced at the base, the anepisternum large, metepisternum narrow like Izleiina, but they differ strongly in the shape of CuA, by having M 2 long, Sc short and free, and by the presence of strong tibial bristles.Published as part of BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, pp. 1-76 in American Museum Novitates 3433 (1) on page 41, DOI: 10.1206/0003-0082(2004)4332.0.CO;2, http://zenodo.org/record/471233