Article by Calvin R. Fremling and D. Kent Johnson regarding mayflies as biological indicators of water quality in areas of the Upper Mississippi River. Fremling and Johnson credited respectively as Biology Department, Winona State University, Winona, Minnesota, 55987, USA, and Metropolitan Waste Control Commission, St. Paul, Minnesota, 55101, USA. Abstract: Hexagenia mayflies are good indicators of general water quality because they have long life cycles and because their burrowing nymphs, which are unable to tolerate anaerobic conditions or swim long distances, live in sediments where toxins tend to accumulate. While chemical tests only describe water quality in terms of specific parameters and times, Hexagenia distribution indicates ...