This contribution presents new evidence on a specific kind of prehistoric sacred landscape, a high-altitude monumental sanctuary characterized by the presence of megalithic stelae decorated with animal relief. The contribution is based on the case-study of Karmir Sar, a recently discovered Chalcolithic and Bronze Age site located on the south slope of Mount Aragats (Armenia), at 2850 m a.s.l. Archaeological investigations demonstrate that, in the ancient South Caucasus, vertical zonality in landscape use conditioned all cultural developments and constituted an integrated system—a chain, each link of which was dominant during particular periods. An important link in this chain were transhumant pastoralists, who each summer moved—as they stil...