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Abbreviations used in this paper: ALT, alternative lengthening of telomeres; APB, ALT-associated PML body; coIP, coimmunoprecipitation; GNL3L, guanine nucleotide-binding protein-like 3-like; HR, homologous recombination; IDF, interstitial damage foci; NS, nucleostemin; PML, promyelocytic leukemia; TA+, telomerase active; TIF, telomere dysfunction-induced foci; TRF, telomeric repeat-binding factor