Proxy evidence and model experiments suggest that the El Niño Southern Oscillation (ENSO) system varied on both orbital and millennial timescales during the late Quaternary. A leading explanation for this variability is a mechanism called the ocean dynamical thermostat, whereby increased radiative energy invigorates atmospheric circulation resulting in cooling in the eastern part of the Pacific Ocean similar to that observed during the La Niña (cold) phase of ENSO. Proxy records of Mg/Ca-derived sea surface temperature (SST) and climate models show an increased temperature gradient across the Pacific corresponding to a more La Niña-like state during the early to mid Holocene, driven by enhanced insolation during Boreal summer/fall. Previous...