Subspecies and Distribution. P. d. dasymallus Temminck, 1825 — Osumi and Tokara Is (Kuchinoerabujima, Kuchinoshima, Nakanojima, Suwanosejima, Tairajima, Akusekijima, Takarajima), N Ryukyu Is, Japan. P. d. daitoensis Kuroda, 1921 — Daito Is (Kita-Daito, Minami-Daitd), E Ryukyu Is, Japan. P. d. formosus Sclater, 1873 — Taiwan (Lutao, Lanyu, and Guishan Dao Is) and Philippines (Batan, Babuyan, Calayan, Dalupiri, Fuga, and Barit Is). Probably extinct on Taiwan mainland. P. d. inopinatus Kuroda, 1933 — Anami and Okinawa Is (Okinawajima, Izenajima, Kourijima, Iejima, Minnajima, Ikeijima, Henzajima, Hamahigajima, Kudakajima, Agunijima, Iheyajima, Yoronjima, Tokasikijima), C Ryukyu Is, Japan. P. d. yayeyamae Kuroda, 1933 — Miyako and Yaeyama Is (Ishigakijima, Iriomotejima, Kohamajima, Hatomajima, Taketomijima, Haterumajima, Kurojima, Yonagunijima, Aragusukujima, Taramajima, Irabujima, Miyakojima, Ikemajima, Ogamijima, Kurimajima, Shimonjijima), S Ryukyu Is, Japan. in Pteropodidae

Subspecies and Distribution. P. d. dasymallus Temminck, 1825 — Osumi and Tokara Is (Kuchinoerabujima, Kuchinoshima, Nakanojima, Suwanosejima, Tairajima, Akusekijima, Takarajima), N Ryukyu Is, Japan. P. d. daitoensis Kuroda, 1921 — Daito Is (Kita-Daito, Minami-Daitd), E Ryukyu Is, Japan. P. d. formosus Sclater, 1873 — Taiwan (Lutao, Lanyu, and Guishan Dao Is) and Philippines (Batan, Babuyan, Calayan, Dalupiri, Fuga, and Barit Is). Probably extinct on Taiwan mainland. P. d. inopinatus Kuroda, 1933 — Anami and Okinawa Is (Okinawajima, Izenajima, Kourijima, Iejima, Minnajima, Ikeijima, Henzajima, Hamahigajima, Kudakajima, Agunijima, Iheyajima, Yoronjima, Tokasikijima), C Ryukyu Is, Japan. P. d. yayeyamae Kuroda, 1933 — Miyako and Yaeyama Is (Ishigakijima, Iriomotejima, Kohamajima, Hatomajima, Taketomijima, Haterumajima, Kurojima, Yonagunijima, Aragusukujima, Taramajima, Irabujima, Miyakojima, Ikemajima, Ogamijima, Kurimajima, Shimonjijima), S Ryukyu Is, Japan.Published as part of Don E. Wilson & Russell A. Mittermeier, 2019, Pteropodidae, pp. 16-162 in Handbook of the Mammals of the World – Volume 9 Bats, Barcelona :Lynx Edicions on page 155, DOI: 10.5281/zenodo.644881