Stalobrachys alboapicata Constant 2018, gen. et comb. nov.

Stalobrachys alboapicata (Jacobi, 1928) gen. et comb. nov. Figs 3, 50–54 Olonia alboapicata Jacobi, 1928: 5 (described). Olonia alboapicata – Metcalf 1956: 65 (catalogued). Diagnosis As for genus. Etymology The species is epithet is formed from albus (adjective, Latin) = ʻwhiteʼ and ʻapexʼ. It refers to the white apical patch of the posterior wing. Material examined Holotype AUSTRALIA • ♀; N Queensland, Cedar Creek; [coordinates of Ravenshoe: 17°36′33″ S, 145°29′01″ E]; “Queensl. Mjöberg”, “Cedar creek”, “ Olonia alboapicata Jac., A. Jacobi determ.”, “Typus”, “Typus”; NHRS (Fig. 52). Notes Jacobi (1928) erroneously stated that the type specimen is a male. He was apparently prone to confusing males and females of Eurybrachidae, as he made the same mistake for the types of Elthenus modestus Jacobi, 1928 and Olonia nigroapicata Jacobi, 1928 (Constant 2005b, 2006b, respectively). Cedar Creek was a Jirrbal aboriginal camp located at the southern edge of Ravenshoe (North Queensland). It covers 38 ha of grassland and woodland on a basalt plateau bordered by North Cedar Creek, South Cedar Creek and the Millstream River in the west. It represented an Eucalyptus “pocket” at the edge of the rainforest. Mjöberg and his staff sampled the location 29 Mar.–17 Apr. 1913 (Ferrier 2006, 2015). Additional material AUSTRALIA • 1 ♂; N Queensland, Ravenshoe; 17°36′33″ S, 145°29′01″ E; Jan. 1921; USNM. Description MEASUREMENTS AND RATIOS. LT: ♂ (n = 1): 8.8 mm; ♀ (n = 1): 10.0 mm; BV/LV = 3.9; BF/LF = 2.0; LP+LM/BT = 0.61; Ltg/BTg = 2.17; LW/BW = 1.6. Male HEAD (Fig. 53A–D). Vertex concave, with anterior and posterior margins parallel, curved, slightly wrinkled longitudinally; yellowish brown with an impression marked with darker colour at each side of disc. Frons dark brown variegated with blackish. Clypeus elongate, coloured as frons. Genae brown. Labium brown, reaching metacoxae. Antennae dark brown; scape short, ring-shaped; pedicel subcylindrical, slightly narrowing towards apex. THORAX (Fig. 53A–C). Pronotum yellowish brown, dark brown at lateral angles; slightly wrinkled; median carina obsolete and 2 small, impressed points on disc. Lateral fields of prothorax dark brown. Mesonotum dark brown, transversely wrinkled; small yellowish spot at apex of scutellum; median and peridiscal carinae weakly marked. Red ventrally. Tegulae brown. TEGMINA (Fig. 53A, C). Brown, darker on clavus and apicocostal half of membrane, irregularly marked with small yellowish spots; one larger pale yellowish marking on vein A1 at midlength of clavus; triangular white marking on costal margin after nodal line; white marking at apicosutural angle. Costal and sutural margins subparallel until nodal line, then slightly converging; costal margin slightly sinuate; apical margin obliquely rounded. POSTERIOR WINGS (Fig. 53E). Dark brown, paler on anal area and becoming darker, nearly black on about distal third; elongate, transverse, subtriangular white marking at apicocostal angle, extending on 4 cells. Margin of anal area slightly sinuate; sutural margin with 2 clefts, cubital one not marked. LEGS (Fig. 53A–D). Pro- and mesocoxae reddish brown. Pro- and mesofemora dark brown with reddish and yellowish spots marking obsolete rings. Pro- and mesotibiae dark brown with yellowish markings on 2 obsolete rings. Pro- and mesotarsi dark brown, with basal half of third tarsomere reddish. Metacoxae reddish; metafemora reddish with apex brown. Metatibiae brown, with 3 lateral spines yellowish basally and black distally, and 9 apical black-brown spines. Metatarsi reddish brown, with first tarsomere darker distally and with a ventral group of 9 black spines in two irregular rows. ABDOMEN. Bright red with genital segments yellowish white. MALE GENITALIA (Fig. 54). Pygofer, anal tube and gonostyli whitish. Pygofer higher than long, with basal margin sinuate in lateral view; in lateral view, dorsal 2/5 very narrow and ventral 2/5 more than two times as broad as dorsal portion; strong posterior projection at half heigth, about twice as long as ventral portion (Fig. 54A, C). Anal tube dorsoventrally flattened, elongate, rounded apically in dorsal view and with posteromedian portion slightly depressed; epiproct at basal ½ (Fig. 54A–B). Gonostyli separated, large, mostly dorsoventally flattened, with elongate basidorsal process directed dorsally and bearing spoon-shaped process apically. Aedeagus simple; phallobase with dorsal process elongate, projecting posteriorly and with apical part incurving, and ventral part elongate and dorsoventrally flattened and notched apically; phallus membranous, dorsoventrally flattened and bifid on distal ½. Female Similar to male, but with distal cells of posterior wings white instead of having a white spot at the apicosutural angle (Fig. 52). Distribution and biology This species is recorded only from a small area around Ravenshoe, at the limit between the Queensland Tropical Rainforests and the Einasleigh Upland Savanna bioregions (Fig. 3). The specimens were found in January and April, one of them in a habitat of mixed grassland and woodland on a basalt plateau described as a Eucalyptus patch at the edge of the rainforest (Ferrier 2006, 2015). However, host plants remain undocumented.Published as part of Constant, Jérôme, 2018, Revision of the Eurybrachidae XIV. The Australian genera Olonia Stål, 1862 and Stalobrachys gen. nov. (Hemiptera: Fulgoromorpha), pp. 1-97 in European Journal of Taxonomy 486 on pages 83-88, DOI: 10.5852/ejt.2018.486, http://zenodo.org/record/227015