Distribution of ground beetles of the genus Bembidion (Coleoptera, Carabidae) in the agricultural landscape in Northwestern Russia

We have observed seven species of predator beetles of the genus Bembidion in the agricultural landscape of the Leningrad Region (Northwestern Russia) between the years 2008 and 2018. These species reach their highest abundance in sun-exposed areas, especially mesophilic B. quadrimaculatum Linnaeus, 1761, B. properans (Stephens, 1828), B. lampros (Herbst, 1784), and B. femoratum Sturm, 1825. A few mesohygrophilic B. guttula (Fabricius, 1792) and B. gilvipes (Sturm, 1825) live primarily in areas of dense vegetation. Finally, the hygrophilic B. bruxellense (Wesmael, 1835) was only observed on the most humid soils. Assemblages of Bembidion ground beetles were separated in the fields, field boundaries, and adjacent habitats