A revalidation and a new species in the genus Phanaeus (Coleoptera: Scarabaeoidea: Scarabaeidae: Scarabaeinae)

Abstract. The morphology of the dung beetle populations previously known as Phanaeus quadridens (Say, 1835) is studied. As a consequence, Phanaeus arnaudi Nogueira, Moctezuma and Halffter, sp. nov. is described from Mexico, and Phanaeus borealis Olsoufieff, 1924, stat. rev. is resurrected from previous synonymy to full species status, due the lack of evidence to support the idea that body color varies intrapopulationary and conform to a continuous north-south cline. An identification key to the P. quadridens species group is update