Data from: Revision of the early crocodylomorph Trialestes romeri (Archosauria, Suchia) from the lower Upper Triassic Ischigualasto Formation of Argentina: one of the oldest-known crocodylomorphs

Trialestes romeri (Reig) is an early crocodylomorph from the Ischigualasto Formation (late Carnian – early Norian; Ischigualasto – Villa Unión Basin, Argentina) and one of the oldest-known members of this clade. Two specimens of this species are known, the holotype (PVL 2561) and a referred specimen (PVL 3889), both consisting of associated cranial and postcranial remains. These specimens are incomplete and poorly preserved thus leading previous authors to propose different phylogenetic hypotheses for the species. Trialestes romeri was originally interpreted as a basal dinosaur, subsequently considered to be a crocodylomorph with some dinosaurian characters (e.g. mesotarsal ankle joint, functionally tridactyl pes), and even proposed as a chimera representing two different genera. Some recent workers have mentioned and discussed Trialestes, but none have described it in detail or included it in a quantitative phylogenetic analysis. Here, we describe in detail all the material assignable to the species and test its phylogenetic relationships using a comprehensive data matrix focused on early archosaurs. We support the referral of PVL 3889 to Trialestes and reject the presence of a mesotarsal ankle joint in this specimen. We recovered Trialestes within Crocodylomorpha, closer to Crocodyliformes than Pseudhesperosuchus, Hesperosuchus, Dromicosuchus and Sphenosuchus. Several stratigraphical units worldwide approximately coeval with the Ischigualasto Formation, have also yielded early crocodylomorph remains, such as the upper Santa María (Brazil), Pekin (USA), and lower Maleri (India) formations. However, the crocodylomorph specimens recovered from those units are more incomplete, possess uncertain phylogenetic affinities, or are part of chimaerae. Therefore, Trialestes represents the most completely known of the earliest non-crocodyliform crocodylomorph taxa known to date.Additional Supporting InformationThis file contains four appendices: the character list (Appendix S1), the character scorings modified from the original data matrix of Zanno et al. (2015) for Carnufex carolinensis (Appendix S2), the table of specimens used to build the palaeomap of Triassic crocodylomorphs (Fig. 15) (Appendix S3), and the references of the Additional Supporting Information (Appendix S4).Lecuona et al 2016 - Additional Sup. Information.pdfData matrix used for the analysisThis data matrix is based upon a modified version of the data matrix of Lecuona (2013), which in turn, was modified from the analysis of Nesbitt (2011). The changes proposed by Butler et al. (2014) were also in this data matrix (also Lecuona et al. unpub. data). Finally, we added the early crocodylomorph Carnufex carolinensis (Zanno et al. 2015, Drymala and Zanno 2016), modified scorings for several taxa, and modified the formulation of the characters 219 and 241. Three terminals were included for Trialestes, the holotype (PVL 2561), the referred specimen (PVL 3889), and the combination of both. The final data matrix consists on 87 terminals and 483 characters.Lecuona et al 2016 - Data matrix of the analysis.ne