Data from: Palaeobiogeography and evolutionary patterns of the larger foraminifer Borelis de Montfort (Borelidae)

The palaeobiogeography of the alveolinoid Borelis species reveals the evolutionary patterns leading to the two extant representatives, which occur in shallow-water tropical carbonate, coral reef-related settings. Type material and new material of fossil Borelis species, along with Recent specimens were studied to assess their taxonomic status, species circumscriptions (based on proloculus size, occurrence of Y-shaped septula, and the index of elongation), palaeobiogeography and evolutionary dynamics. The species dealt with here are known from exclusively fossil (B. pygmaea, B. inflata, B. philippinensis, B. melo, B. curdica), and from fossil and modern (B. pulchra, B. schlumbergeri) specimens. For the first time, fossil and Recent Borelis specimens are illustrated via micro-computed tomography scanning images. Depending on the occurrence of Y-shaped septula, two lineages are distinguished. Deriving from the middle–upper Eocene Borelis. vonderschmitti, the first lineage includes B. inflata, B. pulchra and B. pygmaea, lacking Y-shaped septula. The first species bearing Y-shaped septula is the Rupelian B. philippinensis of the western Indo-Pacific. The westward migrants of B. philippinensis into the Mediterranean gave rise to B. melo (Aquitanian–Messinian) and B. curdica (Burdigalian–Tortonian). These two species became isolated from the Indo-Pacific by the Langhian eastern closure of the Mediterranean basin and disappeared during the Messinian Salinity Crisis. Since the Tortonian, B. schlumbergeri, which descended from B. philippinensis, has inhabited the Indo-Pacific along with B. pulchra. From the central Pacific Ocean, B. pulchra reached the Caribbean area before the early Piacenzian closure of the Central America seaway.Video model of Borelis curdica (Reichel, 1937), early Langhian, Hungary This is the video model of the X-ray Computed Tomography (CT)-scanning images of Borelis curdica (Reichel, 1937) from the early Langhian, Bagoly Hill, Letkés village, Hungary. This specimen is illustrated in Fig. 10 of the paper. Ungheria1_70kvp_100uA_1s_300_300_249_substack.avi inverted_Ungheria1_70kvp_100uA_1s_300_300_249_substack.avi Image stack of computed tomography of Borelis curdica Image stack of the X-ray micro-tomography (micro-CT) scan of the fossil larger foraminifer Borelis curdica (Reichel, 1937). This specimen is illustrated in the Fig. 10 of the paper. The text file contains scan details and settings. Open this file via ImageJ software. Ungheria1_70kvp_100uA_1s_300_300_249_substack.ti