Strongylophthalmyia immaculata Hennig

Strongylophthalmyia immaculata Hennig (Figs. 23, 54, 65) Strongylophthalmyia immaculata Hennig 1940: 309; 1941a: 136. Frey 1956: 129. Steyskal 1971: 144; 1977: 22. Evenhuis 1989: 472. Yang & Wang 1998: 459. Papp 2005: 195. Iwasa & Evenhuis 2014: 103. Diagnosis. Strongylophthalmyia immaculata is similar in appearance to S. palpalis and S. sumatrana, n. sp. on the basis of the yellowish to reddish brown thorax, but it can easily be distinguished from those two by the nonmodified male palpus (bacilliform in S. immaculata; flared apically in S. sumatrana, n. sp.; spherical in S. palpalis). Redescription. Lengths [from single examined male]. Ƌ: body, 2.2 mm; wing, 2.0 mm. Male. Head: globular, frons shining brown above ocellar tubercle, yellow below ocellar tubercle to ptilinal suture; face yellowish white; gena yellowish white, silvery pollinose, row of short white hairs ventrally; occiput brown, yellow posteroventrally; clypeus thin, white; palpus bacilliform, grayish brown, with white hairs ventrally, single long white seta apically; proboscis brown. Antenna (Fig. 23) yellow; flagellomere subspherical, clothed with white hairs, with barely perceptible minute dorsal process located just anterior to base of arista; arista two times length of flagellomere, styliform, bare. Thorax: shining; mesonotum and scutellum brown dorsally, mesonotum yellowish laterally, with numerous brown short hair-like setulae; pleura yellow, katepisternum with white hairs ventrally near mid coxa. Wing: hyaline; vein R2+3 nearly straight, ending in costa at level of crossvein dm-cu; crossvein r-m at basal onethird of cell dm; veins R4+5 and M1+2 slightly converging distally; halter white. WIP (Fig. 54). Generally brassy colored throughout; a thin band of magenta along costa in cell r2+3; cell r1 greenish; anal lobe with subbasal spot of blue surrounded by thin band of magenta. Legs: yellow; fore coxa with 1–2 long yellowish white hairs; fore femur (Fig. 65) dorsally with 6 short to minute black thorn-like spicules in wavy row, ventrally with subbasal thorn-like cluster of stiff yellowish setae; hind femur with minute knob-like protuberance basomesally; tarsi white. Abdomen: tergites I–II yellowish to tan, weakly sclerotized medially and laterally; tergites III–VI dark brown, with short sparse yellowish hairs, darker yellow to brown hairs laterally; sternites brown. Male genitalia. Not dissected; epandrium and surstylus shining black, with white hairs; cerci light brown, narrow basally, flared and rounded apically, with long white hairs. Material examined: TAIWAN: 1♂, 25 km east of Chiayi [23.456910°N, 120.765077°E], 700 m, 9 Oct 1993, A. Freidberg (TAU). Remarks. The holotype male and 10 paratype females were described by Hennig (1940) from Toa Tsui Kutsu [= Dongya Cui Da; 22.901815°N, 121.196501°E] and collected along with S. punctata, S. trifasciata, and S. crinita. The holotype male was not available during this study, but a specimen in TAU determined by Shatalkin as S. immaculata matches the salient characters in Hennig’s description so I am confident of its placement in this subgroup. Although Hennig (1940) described 10 paratype females of S. immaculata, unless they were collected in copula, there is no sure way of associating females with males of this species. They are tentatively left here as paratypes of S. immaculata, but further study may show them to be females of another species. Distribution: Taiwan.Published as part of Evenhuis, Neal L., 2016, World review of the genus Strongylophthalmyia Heller (Diptera: Strongylophthalmyiidae). Part I: Introduction, morphology, species groups, and review of the Strongylophthalmyia punctata subgroup, pp. 201-243 in Zootaxa 4189 (2) on pages 219-223, DOI: 10.11646/zootaxa.4189.2.1, http://zenodo.org/record/16587