Righiodrilus

Key to species of Righiodrilus (updated and modified from Feijoo & Gil 2010) 1. Spermathecae absent................................................................................. 2 - Spermathecae present................................................................................. 3 2. Male pores in xvii........................................................... R. muinane Feijoo & Gil, 2010 - Male pores with tubercula pubertatis in anterior region of xvii or middle ½ 16 – ½ 17.......... R. venancioi (Righi, 1982) 3. One pair of spermathecae.............................................................................. 4 - More than one pair of spermathecae..................................................................... 5 4. Spermathecae in viii–xix............................................................ R. dithecae (Righi, 1988) - Spermathecae in xix–x..................................................... R. guahibo (Righi & Molina, 1994) 5. Four or six pairs of spermathecae....................................................................... 6 - Two or three pairs of spermathecae...................................................................... 7 6. Six pairs of spermathecae in ix–xi, 2 pairs per segment; clitellum in xvi–xxi; male pores in xvii/xviii................................................................................ R. sucunduris (Righi, Ayres & Bittencourt, 1976) - Four pairs of spermathecae, in xiv–xvii; clitellum in xvi–xxiii; male pores in xviii/xix.................. R. gurupi n. sp. 7. Two pairs of spermathecae............................................................................. 8 - Three pairs of spermathecae............................................................................ 9 8. Spermathecae in 7/8 and 8/9.......................................................................... 10 - Not as above........................................................................................ 11 9. Spermathecae in 7/8, 8/9, 9/10......................................................................... 24 - Spermathecae in vi, vii and viii........................................................... R. viseuensis n. sp. 10. Clitellum in xv–xxi.................................................................................. 12 - Clitellum in another position.......................................................................... 13 11. Spermathecae in 9/10 and 10/11............................................ R. amazonius (Zicsi & Csuzdi, 1999) - Spermathecae in another position.......................................................................16 12. Tubercula pubertatis in 1/2 xix–1/2 xx. Male pores in 19/20................................... R. uete (Righi, 1988) - Tubercula pubertatis in xvii–xix. Male pores in 17/18....................................... R. cigges (Righi, 1970) 13. Male pores in 20/21................................................................................. 14 - Not as above........................................................................................15 14. Clitellum in xvii–xxiii; tubercula pubertatis in xix–xxiii.................................... R. freitasi (Righi, 1971) - Clitellum in xvi–xiv; tubercula pubertatis in 1/2 xix–xxii.................................... R. aioca (Righi, 1975) 15. Male pores in 1/2 xix; clitellum in xvi–xxii; tubercula pubertatis in 1/2 xix–1/2 xxi................. R. tico (Righi, 1982) - Male pores in xvii; clitellum in 1/2 xv–1/2 xxiii; tubercula pubertatis in xvii – xx........... R. andake Feijoo & Gil, 2010 16. Spermathecae in 8/9and 9/10......................................................................... 17 - Spermathecae in ix and x............................................................................. 18 17. Clitellum in xv–xxi...................................................................................19 - Clitellum in 1/2 xvi–1/2 xxii or xvi–xxii............................................... R. arapaco (Righi, 1982) 18. Male pores microscopic............................................................ R. oliveirae (Righi, 1982) - Male pores not microscopic........................................................................... 21 19. Male pores in xvii....................................................................................20 - Male pores in 19/20............................................................... R. mairaro (Righi, 1982) 20. Tubercula pubertatis in xviii–xix..................................................... R. paolettii (Righi, 1984) - Tubercula pubertatis in 1/2 xvi–1/2 xix.......................................... R. lebrijae Celis & Rangel, 2015 21. Clitellum in xv–xxii; tubercula pubertatis present.......................................................... 22 - Clitellum in xvi–xxiii; tubercula pubertatis absent................................................ R. moju n. sp 22. Male pores in xvi/xvii; tubercula pubertatis in 1/2 xvi–1/2 xviii............................................... 23 - Male pores in 17/18; tubercula pubertatis in 1/2 xvii–xx................................... R. ortonae (Righi, 1988) 23. Spermathecae in line with setae b....................................................... R. itajo (Righi, 1971) - Spermathecae in line with setae d............................................... R. fontebonensis (Righi, 1988) 24. Male pores in xviii.................................................................................. 25 - Not as above....................................................................................... 26 25. Clitellum in xv–xxii; tubercula pubertatis in 1/2 xvii–1/2 xx.................................................. 27 - Clitellum and tubercula pubertatis in another position...................................................... 28 26. Clitellum in xv–xxi; tubercula pubertatis in 1/2 xvi–1/2 xix.................................................. 29 - Clitellum in xiv–1/2 xxii; tubercula pubertatis in 1/2 xvii–1/2 xx........................... R. inga Feijoo & Gil, 2010 27. Papillae pubertatis absent in xxi; tubercula as grooves with dark color in xviii, xix–xx... R. tocantinensis pola (Righi, 1984) - Papillae pubertatis present in genital field 1/2 xvii–1/2 xx; papilla midventral in xxi.................................................................................................. R. tocantinensis tocantinensis (Righi, 1972) 28. Clitellum in 1/2 xiii–xxii; tubercula pubertatis in xvii–xx................................. R. marcusae (Righi, 1969) - Clitellum in xv–xxiv; tubercula pubertatis in xvii–xxiii............................. R. schubarti (Righi et al., 1978) 29. Male pores in 16/17.............................................................. R. mucupois (Righi, 1970) - Male pores in 17/18................................................................. R. tingus (Righi, 1971)Published as part of Santos, Bianca Tamires Silva Dos, Bartz, Marie Luise Caroline, Hernández-García, Luis Manuel, Rousseau, Guillaume Xavier, Martins, Marlucia Bonifácio & James, Samuel Wooster, 2017, New earthworm species of Righiodrilus (Clitellata, Glossoscolecidae) from eastern Amazonia, pp. 392-400 in Zootaxa 4242 (2) on pages 399-400, DOI: 10.11646/zootaxa.4242.2.11, http://zenodo.org/record/37647