In fibrous paper materials, an exposure to a variation in moisture content causes changes in the geometrical and mechanical properties. Such changes are strongly affected by the inter-fibre bonds, which are responsible for the transfer of the hygro-mechanical response from one fibre to its neighbours in the network, resulting in sheet-scale deformation. Most models developed in literature assume perfect bonding between fibres. In the 3D reality, there is some flexibility in the bond region, even for the perfectly bonded fibres, because of the possibility of deformation gradients through the fibre thickness. In earlier 2D idealizations, perfectly bonded fibres were assumed, implying full kinematic constraint through the entire thickness of t...