This dissertation investigates literary responses to environmental change in nineteenth-century England. Two tropes, chaos in narrative and the microcosm in lyric poetry, suggest how literary works may have been precursors of ecological science. I argue that literary epistemology in the long nineteenth-century developed precocious theories of the way nature operates based on contingent narrative and microcosm systems. These ideas were adopted as empirical strategies once scientific ecology emerged in the twentieth-century, and both tropes are prominent in twenty-first century ecological science. Ecology appeared late among scientific disciplines partly because it relies on cooperation between reduction and holism: climate change theory,...