Porphyrochroa platypoderis Mendonça, Rafael & Ale-Rocha, 2008, sp. nov.

Porphyrochroa platypoderis sp. nov. (Figs. 13 a–i; 19 c; 20) Diagnosis: P. platypoderis sp. nov. Is closely related to P. elongata Mendonça, Rafael & ale-Rocha, 2007 due to the longitudinally divided sternite 8 in two narrow widely separated plates and by the ejaculatory apodeme extended to the level of the sixth abdominal segment when dissected. It differs from the latter species due to the modified midleg tarsi. Description. Male: body length 3.7 mm; wing length 3.2 mm. Frons and face shiny brown. Frons slightly narrower than width of anterior ocellus, subparallel sided. Face slightly longer than frons, length about half width of frons. Proboscis brown, length equal to height of head. Palpus brown, with one small and almost apical seta. Ocellar tubercule with vitreous light-brown ocellus; two ocellar setae; postocellar setae broken. Antenna wholly ligth-brown with conspicuous pruinescence, more dense on flagellum; scape and pedicel medially verticillate; flagellum about four times longer than scape and pedicel combined. Postcranium shiny brown, with distinct gray pruinescence; postcranium setae broken; one pair of parallel vertical setae; postocellar setae distinct, arranged in complete single series with dorsal FIGURE 13. Porphyrochroa platypoderis sp. nov., holotype male: a, Tergites 6–8, dorsal view; b, Sternites 6–8, dorsal view; c, Terminalia (hypandrium, phallus, cercus and subepandrial sclerite), lateral view; d, Epandrium, lateral view, lateral view; e, Hypandrium, posterior view; f, Ejaculatory apodeme, anterior view; g, Cercus, dorsal view; h, Subepandrial sclerite, ventral view; i, Midleg. Abbreviation: a cerc, anterior cercus; cerc, cercus; ej apod, ejaculatory apodeme; goncx apod, gonocoxal apodeme; hypd, hypandrium; p cerc, posterior cercus; ph, phallus; sbepand scl, subepandrial sclerite; ST, sternite; T, tergite. setae stronger. Gena and postgena with brown setae and sparse gray pruinescence. Thorax shiny brown, with gray pruinescence conspicuous and sparse. Thoracic setae: 3 slender ktg; 4 sctl, larger internal pair; aprn and prepm broken; 1 pprn; 2 acr; 4 dc; 1 spal presut; 1 spal psut; 2 ntpl; 1 pal. Fore and hind legs with coxae, trochanters and basal fifth of femora yellow, remaining segments brown. Hind tibia with long dorsal setae; posterior basal tarsomere with two long dorsal setae. Midleg modified (Fig. 13 i), with coxa, trochanter and femur yellow and tibia and tarsi brown; mid tibia with long setae: 1 anterior, 3 dorsal and 1 ventral; mid tarsi modified with long dorsal setae, basal tarsomere projected posteriorly; tarsomere second long and expanding apically; tarsomere third wide; tarsomere fourth narrow and long; and tarsomere fifth long and narrower that fourth; long setae over tarsomere. All legs with short distinct setae. Wing hyaline with conspicuous pterostigma; anal lobe wide (Fig. 19 c). Halter brown. Abdomen shiny brown, with metallic green luster in diverse light incidence angles. Sternites shiny brown. Tergites 6, 7 and 8 subrectangular (Fig. 13 a). Sternite 8 divided longitudinally in two narrow plates widely separated (Fig. 13 b). Terminalia: cercus elongate in lateral view, with depression in center (Fig. 13 c) and with base wider than apex (Fig. 13g). Hypandrium reduced (Figs. 13 c; e). Ejaculatory apodeme greatly elongate, reaching level of sixth abdominal segment when dissected (Fig. 13 c); anterior view represented as in Figure 13 f. Epandrium rectangular with short anteroventral projection and dark anteriorly (Fig. 13 d). Subepandrial sclerite with base and apex with proportional sizes (Fig. 13 h). Female: Unknown. Distribution: BRAZIL (Amazonas). Type material: Holotype male: BRAZIL, Amazonas, Manaus, PDBFF-WWF, Proj.: Bert Klein / xii. 1985, Malaise / Reserva 1301 / Holotype macho Porphyrochroa platypoderis ( INPA ). Paratypes: BRA- ZIL, Amazonas, Manaus, PDBFF-WWF, Proj.: Bert Klein, iv. 1986, Malaise, Reserva 1113 (1 male, MZUSP); v. 1986 (1 male, INPA); i. 1985, Malaise, Reserva 1301 (1 male, MPEG); ii. 1985 (1 male, INPA); iii. 1985 (1 male, INPA); iv. 1985 (1 male, INPA); v. 1985 (1 male, INPA); vi. 1985 (1 male, INPA); viii. 1985 (1 male, INPA); xii. 1985 (1 male, INPA); Novo Aripuanã, 05° 15 ’ 53 ”S, 60 °07’08”W, R. Soka 28.iv–05.vi. 1999, Cols. Ferreira, R. L.; Rocha, R. A., Vidal, J. & Leite, R. S., Suspensa alta – área aberta (1 male, INPA); Holotype condition: postocellar setae, postcranial, aprn and prepm broken. Left wing mounted on microslide in Canada balsam; abdomen and midleg within microvial containing glycerin. Microslide and microvial pinned attached to the specimen. Variation: one paratype with two proclinate setae between the inferior ocellus and the insertion of the antenna, in different heights. Etymology: from Grego p laty = flat, pous = foot. It makes allusion to the presence of midleg flattened structures.Published as part of Mendonça, Mirian Nascimento, Rafael, José Albertino & Ale-Rocha, Rosaly, 2008, Revision of the Brazilian Amazon Basin species of Porphyrochroa Melander (Diptera: Empididae), pp. 1-39 in Zootaxa 1859 on pages 25-27, DOI: 10.5281/zenodo.18363