Acknowledgements: We thank Maximilian Tschol, Lana Dunan and Justin M. J. Travis for useful discussions and comments on the initial results. All simulations were performed using the Maxwell computing cluster at the University of Aberdeen. APC was supported by the University of Aberdeen. GB was supported by a Royal Society University Research Fellowship (UF160614). JMR was supported by NTNU and the Centre for Biodiversity Dynamics (NFR grant 223257).Peer reviewedPublisher PD
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