For nearly one-and-a-half centuries, biologists interested in evolution have been haunted by the question of whether their conceptions are or are not 'Darwinian'. While it may not be unique, this persistent positioning of new developments in relation to a single, pioneering figure is quite exceptional in the history of modern natural science. Physicists currently working in the domains of relativity or quantum theory may refer sometimes to Einstein or Bohr; but their debates are not massively structured by this reference as evolutionary theory has been and remains structured by reference to Darwin. A proximate cause of Darwin's enduring presence is that evolutionary biologists have never stopped reading him. The remarkably numerous editions...