We initiate a new approach to the study of the combinatorics of several parametrizations of canonical bases. In this work we deal with Lie algebras of type A. Using geometric objects called rhombic tilings we derive a Crossing Formula for the action of the crystal operators on Lusztig data for an arbitrary reduced word of the longest Weyl group element. We provide the following three applications of this result. Using the tropical Chamber Ansatz of Berenstein-Fomin-Zelevinsky we prove an enhanced version of the Anderson-Mirkovic conjecture for the crystal structure on MV polytopes. We establish a duality between Kashiwara's string and Lusztig's parametrization, revealing that each of them is controlled by the crystal structure of the other....