In-place operculum demonstrates that the Middle Cambrian Protowenella is a hyolith and not a mollusc

Protowenella is an early–middle Cambrian, isostrophically coiled, calcareous microfossil originally described from the middle Cambrian (Miaolingian) of Australia; it has been referred previously to the molluscan classes Monoplacophora, Helcionelloida or Gastropoda. A unique specimen from the Henson Gletscher Formation (Miaolingian Series, Wuliuan Stage) of North Greenland has a bilaterally symmetrical operculum preserved in place within the shell aperture. Paired cardinal processes and clavicles on the inner side of the operculum indicate that Protowenella was a hyolith morphologically close to the orthothecid Conotheca. Protowenella is transferred from Mollusca to Hyolitha, Order Orthothecida, Family Protowenellidae nov., representing a novel morphological departure from the generally slender cones of other hyoliths