Models inconsistent with altruism cannot explain the evolution of human cooperation

First paragraph: The article "Intuition, deliberation, and the evolution of cooperation," by Bear and Rand (1), uses game theoretic models to examine the role of intuition and deliberation in human cooperation. The premise is that dual processes characterize human social decision making: "(i) automatic, intuitive processes that are relatively effortless but inflexible; and (ii) controlled, deliberative processes that are relatively effortful but flexible" (1). The objective is to "provide a formal theoretical framework for considering the question of whether prosociality is intuitive or whether it requires self-control," and the article concludes that "evolution never favors strategies for which deliberation increases cooperation" (1). However, the evolutionary model suffers from a serious shortcoming; it precludes the survival of altruistic individuals—thought to represent a major share of human populations (2). It is therefore not suitable for addressing whether human cooperative behavior is intuitive