Despite continuing interest in whether plant residues and microwear can give an archaeological 'signature' for the grinding of grain, few studies have looked at what is actually present on ethnographic seed-grinders. In this paper, we map the distribution of use-polish and residues on a classic Central Australian millstone. We begin by setting out an explanatory 'cradle�to�grave' model of the dynamics of millstones to assist our interpretations. We then apply various methods of functional analysis to map the distribution of microwear and residues across the millstone. Several dynamics are evident: (1) the kinetics of grinding create spatial variability in use�polish and residues; (2) systemic factors lead to a palimpsest of different residu...