A pencil of Enriques surfaces with non-algebraic integral Hodge classes

We prove that there exists a pencil of Enriques surfaces defined over Q with non-algebraic integral Hodge classes of non-torsion type. This gives the first example of a threefold with the trivial Chow group of zero-cycles on which the integral Hodge conjecture fails. As an application, we construct a fourfold which gives the negative answer to a classical question of Murre on the universality of the Abel-Jacobi maps in codimension three