Two new genera in the Omphalodes group (Cynoglosseae, Boraginaceae)

Omphalodes (Boraginaceae, Cynoglosseae) phylogenetic relationships are surveyed in the context of the tribe Cynoglosseae, focusing both in the internal relationships among representatives of Omphalodes main subgroups (and including Omphalodes verna, the type species), and their relationships with other Cynoglosseae genera that have been related to the Omphalodes group. Our phylogenetic analysis of ITS and trnL-trnF molecular markers establish close relationships of the American Omphalodes with the genus Mimophytum, and also with Cynoglossum paniculatum and Myosotidium hortensia. The southwestern European annual Omphalodes species form a discrete group deserving taxonomic recognition. We describe two new genera to avoid a paraphyletic Omphalodes genus, accommodating the European annual species in Iberodes and Cynoglossum paniculatum in Mapuchea.Omphalodes (Boraginaceae, Cynoglosseae) molecular phylogenetic relationships are surveyed in the context of the tribe Cynoglosseae, being confirmed that genus Omphalodes is paraphyletic. Our work is focused both in the internal relationships among representatives of Omphalodes main subgroups (and including Omphalodes verna, the type species), and their relationships with other Cynoglosseae genera that have been related to the Omphalodes group. Our phylogenetic analysis of ITS and trnL-trnF molecular markers establish close relationships of the American Omphalodes with the genus Mimophytum, and also with Cynoglossum paniculatum and Myosotidium hortensia. The southwestern European annual Omphalodes species form a discrete group deserving taxonomic recognition. We describe two new genera to reduce the paraphyly in the genus Omphalodes, accommodating the European annual species in Iberodes and Cynoglossum paniculatum in Mapuchea. The pollen of the former taxon is described in detail for the first time