Monogeneans from Pangasiidae (Siluriformes) in Southeast Asia: II. Four new species of

The examination of gills from two rare endemic species of Pangasius Valenciennes, 1840 (Siluriformes, Pangasiidae): P. humeralis Roberts, 1989 (five specimens) and P. lithostoma Roberts, 1989 (six specimens) from the Kapuas River (West Kalimantan province, Borneo Island, Indonesia) revealed the presence of four new species of Monogenea belonging to the genus Thaparocleidus Jain, 1952 (Monogenea, Ancylodiscoidinae) on P. humeralis : T. sinespinae n. sp., T. brevicochleus n. sp., T. kapuasensis n. sp. and T. gustianoi n. sp., and not any on P. lithostoma