A revision of the Western Palaearctic species of Myrmecopora Saulcy, 1864, sensu lato and Eccoptoglossa Luze, 1904 (Coleoptera: Staphylinidae, Aleocharinae, Falagriini).

Die systematische Stellung der Falagriinengattungen Myrmecopora Saulcy, 1864 sensu lato und Eccoptoglossa Luze, 1904, einschließlich der schon beschriebenen Untergattungen und Arten, wird unter Berücksichtigung historischer Entwicklungen untersucht und erörtert. Beide Gattungen werden unter besonderer Berücksichtigung phylogenetischer Aspekte redeskribiert und redefiniert. Folgende Taxa werden phylogenetisch begründet als Untergattungen von Myrmecopora erkannt: Myrmecopora Saulcy s. str., Iliusa Mulsant & Rey, 1874, Xenusa Mulsant & Rey, 1874, Paraxenusa subg. n. und Euphorbiusa subg. n. Eine Revision der Typen und weiteren westpaläarktischen Materials aus beiden Gattungen ergab insgesamt 22 valide Myrmecopora- sowie 2 Eccoptoglossa-Arten, darunter 6 neue Taxa: Myrmecopora (s. str.) wunderlei sp. n. aus Südanatolien, M. (s. str.) convexula sp. n. aus Westanatolien und Südostbulgarien, M. (s. str.) elisa sp. n., M. (s. str.) plana sp. n., M. (s. str.) fornicata sp. n., die letzteren drei von Kreta, sowie M. (Xenusa) oweni sp. n. aus Südengland, Nord- und Westfrankreich. Myrmecopora turanica Eppelsheim, 1888, wird in die Gattung Eccoptoglossa gestellt. Folgende Synonymisierungen werden vorgenommen: M. (Iliusa) fugax (Erichson, 1839) = M. uludaghensis Fagel, 1971, syn. n., M. (Xenusa) uvida (Erichson, 1840) = M. meridiogallica Scheerpeltz, 1972, syn. n., M. (Xenusa) boehmi Bernhauer, 1910 = M. sydowi Bernhauer, 1927, syn. n., = Xenusa mediterranea Fagel, 1970, syn. n., M. (Xenusa) sulcata (Kiesenwetter, 1850) = Xenusa carica Fagel, 1970, syn. n., M. (Xenusa) simillima (Wollaston, 1864) = M. lohmanderi Bernhauer, 1927, syn. n., M. (Xenusa) minima Bernhauer, 1900 = M. buresi Rambousek, 1910, syn. n., = Xenusa pamphylica Fagel, 1969, syn. n., Eccoptoglossa obscura Luze, 1904 = Mymecopora afghanica Scheerpeltz, 1972, syn. n. Myrmecopora (s. str.) pygmaea (Sachse, 1852), von früheren Autoren als Synonym von M. (s. str.) crassiuscula (Aubé, 1850) betrachtet, wird revalidisiert. Für Myrmecopora publicana Saulcy, Myrmedonia pygmaea Sachse, Tachyusa laesa Erichson, T. uvida Erichson, T. sulcata Kiesenwetter, T. simillima Wollaston, T. maritima Wollaston, Myrmecopora sydowi Bernhauer, M. boehmi Bernhauer, M. minima Bernhauer, M. lohmanderi Bernhauer, M. bernhaueri Koch, M. brevipes Butler, M. gravata Peyerimhoff, M. turanica Eppelsheim und Eccoptoglossa obscura Luze werden Lectotypen, für Tachyusa fugax Erichson ein Neotypus designiert. Die Beschreibungen der behandelten Gattungen, Untergattungen und Arten werden durch Abbildungen relevanter ektoskelettaler Merkmale und der Genitalien, durch ausführliche Bestimmungsschlüssel sowie durch Angaben zur Verbreitung und Bionomie ergänzt.StichwörterColeoptera, Staphylinidae, Aleocharinae, Falagriini, Myrmecopora, Xenusa, Iliusa, Eccoptoglossa, Palaearctic, Europe, distribution, ecology, taxonomy, revision, new subgenera, new species, lectotype designation, neotype, designation, new synonyms.Nomenklatorische Handlungenobscura Luze, 1904 (Eccoptoglossa), Lectotypeturanica (Eppelsheim, 1888) (Eccoptoglossa), Lectotype; comb. n. hitherto Myrmecopora (Ilyusa) turanicaEuphorbiusa Assing, 1997 (Myrmecopora), sgen. n.lohmanderi Bernhauer, 1927 (Myrmecopora), Lectotype; syn. n. of Myrmecopora (Xenusa) simillima (Wollaston, 1864)Paraxenusa Assing, 1997 (Myrmecopora), sgen. n.sydowi Bernhauer, 1927 (Myrmecopora), syn. n. of Myrmecopora (Xenusa) boehmi Bernhauer, 1910uludaghensis Fagel, 1971 (Myrmecopora), syn. n. of Myrmecopora (Iliusa) fugax (Erichson, 1839)gravata Peyerimhoff, 1945 (Myrmecopora (Euphorbiusa)), Lectotypefugax (Erichson, 1839) (Myrmecopora (Iliusa)), Neotype described as Tachyusa fugaxafghanica Scheerpeltz, 1972 (Myrmecopora (Ilyusa)), syn. n. of Eccoptoglossa obscura Luze, 1904convexula Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.elisa Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.fornicata Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.plana Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.publicana Saulcy, 1864 (Myrmecopora (Myrmecopora)), Lectotypepygmaea (Sachse, 1852) (Myrmecopora (Myrmecopora)), Lectotype described as Myrmedonia pygmaeawunderlei Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.laesa (Erichson, 1839) (Myrmecopora (Paraxenusa)), Lectotype, comb. n. hitherto Tachyusa laesaanatolica (Fagel, 1969) (Myrmecopora (Xenusa)), comb. n. hitherto Xenusa anatolicabernhaueri Koch, 1936 (Myrmecopora (Xenusa)), Lectotypeboehmi Bernhauer, 1910 (Myrmecopora (Xenusa)), Lectotypeburesi Rambousek, 1910 (Myrmecopora (Xenusa)), syn. n. of Myrmecopora (Xenusa) minima Bernhauer, 1900maritima (Wollaston, 1860) (Myrmecopora (Xenusa)), Lectotype described as Tachyusa maritimameridiogallica Scheerpeltz, 1972 (Myrmecopora (Xenusa)), syn. n. of Myrmecopora (Xenusa) uvida (Erichson, 1840)minima Bernhauer, 1900 (Myrmecopora (Xenusa)), Lectotypeoweni Assing, 1997 (Myrmecopora (Xenusa)), spec. n.simillima (Wollaston, 1864) (Myrmecopora (Xenusa)), Lectotype described as Tachyusa simillimasulcata (Kiesenwetter, 1850) (Myrmecopora (Xenusa)), Lectotype described as Tachyusa sulcatauvida (Erichson, 1840) (Myrmecopora (Xenusa)), Lectotype described as Tachyusa uvidacarica Fagel, 1970 (Xenusa), syn. n. of Myrmecopora (Xenusa) sulcata (Kiesenwetter, 1850)mediterranea Fagel, 1970 (Xenusa), syn. n. of Myrmecopora (Xenusa) boehmi Bernhauer, 1910pamphylica Fagel, 1969 (Xenusa), syn. n. of Myrmecopora (Xenusa) minima Bernhauer, 1900The systematic position and taxonomic history of the falagriine genera Myrmecopora Saulcy, 1864 sensu lato and Eccoptoglossa Luze, 1904, including the subgenera and species described prior to the present study are outlined and discussed. Both genera are redescribed and redefined with special reference to phylogenetic relationships. Based on phylogenetic arguments, the following taxa are recognized as subgenera of Myrmecopora: Myrmecopora Saulcy s. str., Iliusa Mulsant & Rey, 1874, Xenusa Mulsant & Rey, 1874, Paraxenusa subg. n. and Euphorbiusa subg. n. A revision of the types and further Western Palaearctic material of both genera yielded 22 valid species of Myrmecopora and 2 species of Eccoptoglossa, among them six new taxa: Myrmecopora (s. str.) wunderlei sp. n. from Southern Anatolia, M. (s. str.) convexula sp. n. from Western Anatolia and Southeastern Bulgaria, M. (s. str.) elisa sp. n. from Crete, M. (s. str.) plana sp. n. from Crete, M. (s. str.) fornicata sp. n. from Crete and M. (Xenusa) oweni sp. n. from Southern England, Northern and Western France. Myrmecopora turanica Eppelsheim, 1888, is transferred to Eccoptoglossa. The following new synonymies are established: M. (Iliusa) fugax (Erichson, 1839) = M. uludaghensis Fagel, 1971, syn. n., M. (Xenusa) uvida (Erichson, 1840) = M. meridiogallica Scheerpeltz, 1972, syn. n., M. (Xenusa) boehmi Bernhauer, 1910 = M. sydowi Bernhauer, 1927, syn. n., = Xenusa mediterranea Fagel, 1970, syn. n., M. (Xenusa) sulcata (Kiesenwetter, 1850) = Xenusa carica Fagel, 1970, syn. n., M. (Xenusa) simillima (Wollaston, 1864) = M. lohmanderi Bernhauer, 1927, syn. n., M. (Xenusa) minima Bernhauer, 1900 = M. buresi Rambousek, 1910, syn. n., = Xenusa pamphylica Fagel, 1969, syn. n., Eccoptoglossa obscura Luze, 1904 = Mymecopora afghanica Scheerpeltz, 1972, syn. n. Myrmecopora (s. str.) pygmaea (Sachse, 1852), previously treated as a synonym of M. (s. str.) crassiuscula (Aubé, 1850), is regarded as a valid species. Lectotypes are designated for Myrmecopora publicana Saulcy, Myrmedonia pygmaea Sachse, Tachyusa laesa Erichson, T. uvida Erichson, T. sulcata Kiesenwetter, T. simillima Wollaston, T. maritima Wollaston, Myrmecopora sydowi Bernhauer, M. boehmi Bernhauer, M. minima Bernhauer, M. lohmanderi Bernhauer, M. bernhaueri Koch, M. brevipes Butler, M. gravata Peyerimhoff, M. turanica Eppelsheim and Eccoptoglossa obscura Luze. A neotype is designated for Tachyusa fugax Erichson. In addition to the descriptions of the genera, subgenera and species dealt with in the study, drawings of relevant external characters and the genitalia, detailed diagnostic keys as well as data on distribution and bionomics are provided.KeywordsColeoptera, Staphylinidae, Aleocharinae, Falagriini, Myrmecopora, Xenusa, Iliusa, Eccoptoglossa, Palaearctic, Europe, distribution, ecology, taxonomy, revision, new subgenera, new species, lectotype designation, neotype, designation, new synonyms.Nomenclatural Actsobscura Luze, 1904 (Eccoptoglossa), Lectotypeturanica (Eppelsheim, 1888) (Eccoptoglossa), Lectotype; comb. n. hitherto Myrmecopora (Ilyusa) turanicaEuphorbiusa Assing, 1997 (Myrmecopora), sgen. n.lohmanderi Bernhauer, 1927 (Myrmecopora), Lectotype; syn. n. of Myrmecopora (Xenusa) simillima (Wollaston, 1864)Paraxenusa Assing, 1997 (Myrmecopora), sgen. n.sydowi Bernhauer, 1927 (Myrmecopora), syn. n. of Myrmecopora (Xenusa) boehmi Bernhauer, 1910uludaghensis Fagel, 1971 (Myrmecopora), syn. n. of Myrmecopora (Iliusa) fugax (Erichson, 1839)gravata Peyerimhoff, 1945 (Myrmecopora (Euphorbiusa)), Lectotypefugax (Erichson, 1839) (Myrmecopora (Iliusa)), Neotype described as Tachyusa fugaxafghanica Scheerpeltz, 1972 (Myrmecopora (Ilyusa)), syn. n. of Eccoptoglossa obscura Luze, 1904convexula Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.elisa Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.fornicata Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.plana Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.publicana Saulcy, 1864 (Myrmecopora (Myrmecopora)), Lectotypepygmaea (Sachse, 1852) (Myrmecopora (Myrmecopora)), Lectotype described as Myrmedonia pygmaeawunderlei Assing, 1997 (Myrmecopora (Myrmecopora)), spec. n.laesa (Erichson, 1839) (Myrmecopora (Paraxenusa)), Lectotype, comb. n. hitherto Tachyusa laesaanatolica (Fagel, 1969) (Myrmecopora (Xenusa)), comb. n. hitherto Xenusa anatolicabernhaueri Koch, 1936 (Myrmecopora (Xenusa)), Lectotypeboehmi Bernhauer, 1910 (Myrmecopora (Xenusa)), Lectotypeburesi Rambousek, 1910 (Myrmecopora (Xenusa)), syn. n. of Myrmecopora (Xenusa) minima Bernhauer, 1900maritima (Wollaston, 1860) (Myrmecopora (Xenusa)), Lectotype described as Tachyusa maritimameridiogallica Scheerpeltz, 1972 (Myrmecopora (Xenusa)), syn. n. of Myrmecopora (Xenusa) uvida (Erichson, 1840)minima Bernhauer, 1900 (Myrmecopora (Xenusa)), Lectotypeoweni Assing, 1997 (Myrmecopora (Xenusa)), spec. n.simillima (Wollaston, 1864) (Myrmecopora (Xenusa)), Lectotype described as Tachyusa simillimasulcata (Kiesenwetter, 1850) (Myrmecopora (Xenusa)), Lectotype described as Tachyusa sulcatauvida (Erichson, 1840) (Myrmecopora (Xenusa)), Lectotype described as Tachyusa uvidacarica Fagel, 1970 (Xenusa), syn. n. of Myrmecopora (Xenusa) sulcata (Kiesenwetter, 1850)mediterranea Fagel, 1970 (Xenusa), syn. n. of Myrmecopora (Xenusa) boehmi Bernhauer, 1910pamphylica Fagel, 1969 (Xenusa), syn. n. of Myrmecopora (Xenusa) minima Bernhauer, 190