One consequence of the homological mirror symmetry conjecture predicts that many varieties will have ``hidden symmetries" in the form of autoequivalences of their derived categories of coherent sheaves which do not correspond to any automorphism of the underlying variety. In fact the fundamental groupoid of a certain "complexified Kaehler moduli space" conjecturally acts on the derived category. When the space in question is the cotangent bundle of a flag variety, actions of this kind have been studied intensely in the context of geometric representation theory and Kahzdan-Lusztig theory. We establish the conjectured group action on the derived category of any variety which arises as a symplectic or hyperkaehler reduction of a linear repres...