Reduced versions of the matrix information included in the animal models shown in this paper.

(a) The genetic relatedness matrix, derived from a pedigree, which contains the probabilities that individuals i and j share alleles that are identical by descent. (b) The environmental similarity matrix, which contains elements Sn(i,j) (written in the figure as S, as the same equation is used for elements of Sepi) measuring the Euclidean distance between the environment individuals experience, p and q being the scaled environmental vectors for individuals i and j across l environments. (c) Epiallelic similarity, which contains elements Sepi(i,j), measuring the Euclidean distance between individuals’ epigenetic profiles, where p and q are the scaled epigenetic vectors for individuals i and j, across l epigenetic islands. (d) The geodesic distance matrix (social network connectedness) that shows the proximity of individuals in the social network, based on minimum path lengths (gi,j).