Simulation results showing the output reference frame response characteristics of a model incorporating a population of sigmoidal modulated input neurons trained with the modified learning rule 18: Delayed postsynaptic firing rate with anti-Hebbian learning.

The scatter plot shows the reference frame response characteristics of all output neurons before and after training. Conventions as for Fig 14. It is evident that training had the effect of increasing the head-centredness values of most output neurons. Indeed many more head-centred output neurons are present in the trained model than in the untrained model.