Weight vectors of two typical output neurons.

The top row shows the weight vectors of two typical output neurons that develop when the input neurons have peaked eye-position gain modulation and the network is trained with either the Hebbian learning rule (A) or the trace learning rule (B). The bottom row shows the weight vectors of two typical output neurons when the input neurons have monotonic eye-position gain and the network is trained with either the Hebbian learning rule (C) or the standard trace learning rule (D).