Summary of the resulting directionality.

A) Raster plot of the spiking activity of the two networks, black dots indicate the spiking activity of the leader, dots in color, the activity of the follower. B-C) Global PRC obtained from direct perturbation of the leader in black and in the follower in color. In panel B, the perturbation is made on the E-cells, in panel C, the perturbation is made on the I-cells. Lower right panel illustrates the emerging directionality of the signal flow. Parameters: Ne = Ni = 5000; τe = τi = 10; τse = τsi = 1; ; Jee = 0; Jei = 15; Jii = 0; Jie = 15; ; vth = 500; vr = −500; ; Gee = 0.1; Gie = 0.5; d = 6; direct perturbations are made with a square wave current pulse (amplitude 1, duration 0.1).