The Pyrenees represents an orogen that developed diachronously, from east to west, between the Late Cretaceous and Miocene. Here, we use detrital zircon fission-track thermochronological analyses and U-Pb geochronology, interpreted within the context of the thermal and tectono-sedimentary development of the orogen, to construct a 3-stage model for south-central Pyrenean sediment routing system evolution as follows: (1) Late Cretaceous to Paleocene: Oblique convergence and topographic growth initiates in the eastern Pyrenees. After erosion and removal of the “cover layer”, south-central Pyrenean basins are supplied with zircons cooled during the Late Cretaceous (∼78 Ma), with a fission-track lag time of ca. 15 Myr, that record early Pyrenean...