\u3ci\u3eMylanodon rosei\u3c/i\u3e, a New Metacheiromyid (Mammalia: Palaeanodonta) from the Late Tiffanian (Late Paleocene) of Northwestern Wyoming

Mylanodon rosei is a new genus and species of late Paleocene metacheiromyid palaeanodont from a new late Tiffanian locality, Y2K Quarry, in the Clarks Fork Basin, Wyoming. The type is an adult dentary with P4 and a molariform double-rooted M1. This provides the first evidence that molariform teeth were retained in early Metacheiromyidae. A second specimen is a juvenile dentary with a partial P3 and an unerupted P4. This is the first juvenile dentition known for a Paleocene metacheiromyid. The new specimens enable determination of dental homologies. Reduction of teeth in early metacheiromyids took place from back to front, opening the characteristic posterior diastema. Both Mylanodon and Propalaeanodon, a slightly older metacheiromyid, are intermediate morphologically and temporally between the older Tiffanian epoicotheriid Amelotabes and the younger Clarkforkian and Wasatchian metacheiromyid Palaeanodon. Propalaeanodon has a single-rooted M1, a derived characteristic not found in Mylanodon, suggesting that two lineages are involved and Propalaeanodon was not ancestral to Mylanodon