This thesis investigates the forms, functions and behaviour of tone in the phonology, lexicon, morphosyntax and the phonology-grammar interfaces in Ikaan (Benue-Congo, Nigeria). The analysis is based on an annotated audio corpus of recordings from 29 speakers collected during ten months of fieldwork complemented with participant observation and informally collected data. The study demonstrates that tone operates at a wide range of levels of linguistic analysis in Ikaan. As phonemes, tones distinguish meaning in minimal pairs and are subject to phonological rules. As morphemes, tones and tonal melodies bear meaning in inflection, derivation and reduplication. In the syntax, tones mark phrase boundaries. At the phonology-semantics interface, ...