Red squirrels benefit from human settlement, and nest boxes in mature and old spruce forests are more likely to be occupied when spruce cone crop is abundant.

<p>The effect of (A) standardized amount of built areas within buffers of 300 and 2.5 km on red squirrel occurrence in nest boxes and (B) the effect of spruce cone crop on the estimated coefficient of mature spruce forest, i.e. the interaction between cone crop and spruce forest on red squirrel occurrence.