An initially better-adapted lineage (red, α = 5) competes against a more lineage that produces better variation and therefore adapts quicker (green, α = 85).

<p>First row: short pre-adaption (30 generations), but no per-deme extinctions. Second row: long pre-adaptation (500 generations), per-deme extinctions at mean period T = 64. D = 16, ND = 1024.