Behavioural frequencies in relation to trail use.

<p>Total observed events of marking (A, B) and investigation (C, D) by each age sex class, compared to their expected frequency in relation to their presence on trails containing active marking trees. Comparisons during the breeding (A & C) and non-breeding season (B & D). *** indicates <i>p</i><0.001 in subdivided testing, ** <i>p</i><0.01 and * <i>p</i><0.05.