Chromosome connections in meiosis.

<p>Kinetochores attach homologous chromosomes to opposite halves of the spindle. Homologs are held together by chiasmata, in which recombinant chromatids cross each other. Sisters are held together by cohesins and possibly by catenation of centromeric DNA threads, which have been observed in human mitosis. Cohesion is released in two steps: on chromosome arms to resolve chiasmata and separate homologs in the first meiotic division; and around centromeres to separate sisters in the second meiotic division.