The firing rates of the head direction (HD) cells recorded during the testing phase.

<p>The rates are recorded from a model implemented with a uniform distribution of axonal conduction delays for and synapses within the interval from ms to ms. The model was trained with a rotational velocity of /s. Conventions are as for the top-right plots in <a href="http://www.plosone.org/article/info:doi/10.1371/journal.pone.0058330#pone-0058330-g007" target="_blank">Figures 7</a> and <a href="http://www.plosone.org/article/info:doi/10.1371/journal.pone.0058330#pone-0058330-g008" target="_blank">8</a>. It is clear from the plot that the model can learn to perform path integration of head direction when simulated with a range of axonal conduction delays.