Model path error in conditions that involve smooth pursuit eye movements.

<p>The optic flow that appears on the observer retinal during smooth pursuit of a horizontally moving target is identical in the orientation along heading and orientation along Z-axis conditions. The orientation along heading condition is similar to the gaze along heading condition, except the observer tracks a target that moves in the direction <i>opposite</i> of the path curvature. The orientation along Z-axis condition is similar to the Z-axis condition, except the observer tracks a target that moves in the <i>same</i> direction of the path curvature. Similar to human subjects, the model yields low path errors for all the path radius conditions because model gain fields compensate in the direction opposite that of the eye movements. The model increasingly underestimates path curvature in the orientation along Z-axis condition, similar to humans.