Results of the general linear models developed for males.

<p>(a) The relation between ‘male copulation tie rate’ and the first factor of the PCA. High negative values on factor 1 indicate high-ranking males who often led the pack, who were frequently approached within 1 metre by females and who displayed both a high rate of affiliative behaviour and a low rate of aggressive behaviour towards females; (b) The relation between “rate of refusals that males received by females” and the second factor of the PCA. High negative values on factor 2 indicate a high rate of aggressive behaviour displayed by males towards females.