Community-weighted means (CWM ± SE) for traits: a) body length, b) flight ability, c) carnivory and d) adult overwintering in the three agricultural land-use types: grasslands, winter wheat and sugar beet fields for emerging (●) and actively moving (○) ground beetle.

<p>Note that a value of “1” (in panel b, c and d) indicates that all individuals in a community had good flight ability (panel b), were carnivorous (panel c) or overwintered as adults (panel d). Vice versa, a value of “-1” indicates that all individuals in a community had poor flight ability (panel b), were herbivorous/omnivorous (panel c) or overwintered as larvae (panel d). The broken horizontal line at the value zero of the y-axis in panel b, c and d indicate the point where the proportion of the opposed traits are equal.