Coefficient of variation (CoV; standard deviation/mean*100%) of the half-sarcomere lengths during an isometric contraction at muscle length <i>L</i>/<i>L</i>₀ = 1.2, for models (blue line), M^Fv (red line), M_ATI (black solid line) and (black dashed line).

<p>While actin—titin interactions can damp the magnitude of half-sarcomere—length heterogeneities, the force-velocity relation damps the their temporal evolution.