A coupled ecosystem-circulation model of the North Atlantic is used to examine the individual contributions by wind stress and surface heat fluxes to naturally driven interannual-to-decadal variability of air-sea fluxes of CO2 and O2 during 1948–2002. The model results indicate that variations in O2 fluxes are mainly driven by variations in surface heat fluxes in the extratropics (15°N to 70°N), and by wind stress in the tropics (10°S to 15°N). Conversely, variations in simulated CO2 fluxes are predominantly wind-stress driven over the entire model domain (18°S to 70°N); while variability in piston velocity and surface heat fluxes is less important. The simulated uptake of O2 by the North Atlantic amounts to 70 ± 11 Tmol yr-1 to which the s...