Direct and indirect genetic and fine-scale location effects on breeding date in song sparrows

Acknowledgements We thank the Tsawout and Tseycum First Nations for generously allowing us continuous access to Mandarte Island, Rebecca Sardell and Pirmin Nietlisbach for paternity assignments, Alastair Wilson and Jarrod Hadfield for helpful discussions, Ben Sheldon and two anonymous reviewers for their comments and the National Sciences and Engineering Research Council of Canada and the European Research Council for funding.Peer reviewedPostprin