DOI: 10.1017/S0016756803008380 Printed in the United Kingdom The lost Lower Old Red Sandstone of England and Wales: a record

Abstract – Illite crystallinity data from the Silurian slate belts of England andWales indicate anchizone to low epizone metamorphism during the Acadian deformation in late Early Devonian time. This metamorphic grade implies a substantial overburden, now eroded, of Lower Devonian non-marine sediments of the Old Red Sandstone (ORS) magnafacies. A minimum 3.5 km pre-tectonic thickness of ‘lost ’ ORS is estimated in the southern Lake District and comparable thicknesses in North Wales and East Anglia. Tectonically driven subsidence of the underlying Avalonian crust is required to accommodate such thicknesses of non-marine sediment. One proposed mechanism is flexure of the Avalonian footwall during convergence that continued from Iapetus closure in the Silurian untilAcadian cleavage formation in the Emsian. The evidence for this model in the critical area of northwest England is reviewed and found to be unconvincing. An alternative model is developed following a recent suggestion that the Early Devonian was a period not of continued convergence but of orogen-wide sinistral transtension. Transtensional accommodation of the lost ORS is evidenced by Early Devonian extensional faults, by synchronous lamprophyric magmatism, and by compatibility with previously diagnosed sediment provenance patterns. A summary of Siluro-Devonian tectonostratigraphy for Britain south of the Highland Border emphasizes that, unlike the Scottish Highlands, this area was no